Google Scholar Profile

All publications are provided as final PDF and/or pre-prints below. For final published versions, please email me at

Stoffel MA, SE Johnston, JG Pilkington, JM Pemberton (2020) Genetic architecture and lifetime dynamics of inbreeding depression in a wild mammal. Preprint on BioRxiv: doi:/10.1101/2020.05.27.118877v1

Sparks AM, K Watt, R Sinclair, JG Pilkington, JM Pemberton, TN McNeilly, DH Nussey, SE Johnston (2019) The genetic architecture of helminth-specific immune responses in a wild population of Soay sheep (Ovis aries). PLoS Genetics, 15, e1008461

O’Sullivan RJ, T Aykanat, SE Johnston, A Kane, R Poole, G Rogan, PA Prodöhl, CR Primmer, P McGinnity, TE Reed (2019) Evolutionary stasis of a heritable morphological trait in a wild fish population despite apparent directional selection. Ecology and Evolution, 9, 7096-7111.

Mobley K, H Granroth-Wilding, M Ellmen, J-P Vaha, T Aykanat, SE Johnston, P Orell, J Erkinaro, CR Primmer (2019) Home ground advantage: selection against dispersers promotes cryptic local adaptation in wild salmon. Science Advances, 5, eaav1112. Preprint available on BioRxiv,

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Johnston SE, J Huisman, JM Pemberton (2018) A genomic region containing REC8 and RNF212B is associated with individual recombination rate variation in a wild population of red deer (Cervus elaphus). G3: Genes, Genomes, Genetics, doi: 10.1534/g3.118.200063

Stapley J, PGD Feulner, SE Johnston, AW Santure, CM Smadja (2017) Recombination: the good, the bad and the variable. Philosophical Transactions of the Royal Society, 372: 20170279 [Open Access]

Stapley J, PGD Feulner, SE Johnston, AW Santure, CM Smadja (2017) Variation in recombination frequency and distribution across Eukaryotes: patterns and processes. Philosophical Transactions of the Royal Society, 372: 20160455 [Open Access]

Johnston SE, J Huisman, PA Ellis, JM Pemberton (2017) A high-density linkage map reveal sexually-dimorphic recombination landscapes in red deer (Cervus elaphus). G3: Genes, Genomes, Genetics, 7, 2859-2870

Gonen S, M Battagin, SE Johnston, G Gorjanc, JM Hickey (2017) The potential of shifting recombination hotspots on genetic gain in livestock breeding. Genetics Selection Evolution, 49:55

Lenormand T, J Engelstädter, SE Johnston, E Wijnker, CR Haag (2016) Evolutionary mysteries in meiosis. Philosophical Transactions of the Royal Society, 371, 20160001. Preprint available on [bioRxiv]

Battagin M, G Gorjanc, A-M Faux, SE Johnston, JM Hickey (2016) Effect of manipulating recombination rates on response to selection in livestock breeding programs. Genetics Selection Evolution, 48:44 [PDF]

Johnston SE, C Bérénos, J Slate, JM Pemberton (2016) Conserved genetic architecture underlying individual recombination rate variation in a wild population of Soay sheep. Genetics, 203, 583–598 [PDF]

Barson N, T Aykanat, K Hindar, M Baranski, G Bolstad, P Fiske, C Jacq, A Jensen, SE Johnston, S Karlsson, M Kent, T Moen, E Niemelä, T Nome, T Naesje, P Orell, A Romakkaniemi, H Saegrov, K Urdal, J Erkinaro, S Lien, C Primmer (2015). Sex-dependent dominance at a single locus maintains variation in age at maturity in Atlantic salmon, Nature, 528, 405–408 [PDF]

Aykanat T, SE Johnston, P Orell, E Niemelä, J Erkinaro, CR Primmer (2015) Low but significant genetic differentiation underlies biologically meaningful phenotypic divergence in a large Atlantic salmon population. Molecular Ecology, 24, 5158-5174 [PDF]

Johnston SE, P Orell, VL Pritchard, MP Kent, S Lien, E Niemelä, J Erkinaro, CR Primmer (2014) Genome-wide SNP analysis reveals a genetic basis for sea-age variation in a wild population of Atlantic salmon (Salmo salar). Molecular Ecology, 23, 3452–3468 [PDF]

Aykanat T, SE Johnston, D Cotter, TF Cross, R Poole, PA Prodöhl, T Reed, G Rogan, P McGinnity, CR Primmer (2014) Molecular pedigree reconstruction and estimation of evolutionary parameters in a wild Atlantic salmon river system with incomplete sampling: a power analysis. BMC Evolutionary Biology, 14:68 [PDF]

Johnston SE, J Gratten, C Berenos, JG Pilkington, TH Clutton-Brock, JM Pemberton, J Slate (2013) Life history trade-offs at a single locus maintain sexually selected genetic variation. Nature, 502, 93-95 [PDF] [Supp. Info]

Johnston SE, M Lindqvist, E Niemelä, P Orell, J Erkinaro, MP Kent, S Lien, J-P Vähä, A Vasemägi, CR Primmer (2013) Fish scales and SNP chips: SNP genotyping and allele frequency estimation in individual and pooled DNA from historical samples of Atlantic salmon (Salmo salar). BMC Genomics, 14:439 [PDF]

Bruneaux M*, SE Johnston*, G Herczeg, J Merilä, CR Primmer and A Vasemägi (2013). Molecular evolutionary and population genetic analysis of the nine-spined stickleback using a modified RAD tag approach (*as joint first author). Molecular Ecology, 22, 565-582 [PDF]

Johnston SE, JC McEwan, NK Pickering, JW Kijas, D Beraldi, JG Pilkington, JM Pemberton, J Slate (2011) Genome-wide association mapping identifies the genetic basis of discrete and quantitative variation in sexual weaponry in a wild sheep population. Molecular Ecology, 20, 2555-2566 [PDF]

Johnston SE (2010) The genetic basis of sexual weaponry in a wild sheep population. PhD Thesis, University of Sheffield [PDF]

Slate J, AW Santure, PGD Feulner, EA Brown, AD Ball, SE Johnston & J Gratten (2010) Genome mapping in intensively studied wild vertebrate populations. Trends in Genetics, 26, 275-284 [PDF]

Johnston SE, D Beraldi, AF McRae, JM Pemberton & J Slate (2010) Horn type and horn length genes map to the same chromosomal region in Soay sheep. Heredity104, 196–205 [PDF]

Baxter S, SE Johnston & C Jiggins (2009) Butterfly speciation and the distribution of gene effect sizes fixed during adaptation. Heredity, 102, 57-65 [PDF]